short-beak arrowhead, more...
[Sagittaria engelmanniana subsp. brevirostra (Mack. & Bush) Bogin]
Herbs, perennial, to 70 cm; rhizomes absent; stolons present; corms present. Leaves emersed; petiole terete-ridged, to 39 cm; blades sagittate, 5--20 ´ 2--8 cm, basal lobes equal to or less than remainder of blade. Inflorescences racemes or panicles, of 5--12 whorls, emersed, 25--30 ´ 6--15 cm; peduncles to 45 cm; bracts distinct or if connate, then less than ¼ total length, long-acuminate, 10--40 mm, firm, not papillose; fruiting pedicels spreading to ascending, cylindric, 1--2.5 cm. Flowers to 3.5 cm diam.; sepals recurved to spreading, not enclosing flower; filaments cylindric, longer than anthers, glabrous; pistillate pedicellate, without ring of sterile stamens. Fruiting heads (1.2--)1.5--2.5 cm diam.; achenes cuneate-obovoid, without abaxial keel, 2.1--3.1 ´ 1.4--2.2 mm, beaked; faces --, wings 0--2, ± entire, glands absent; beak lateral, ascending apically, 0.4--1.7 mm. missing datum is whether faces are tuberculate or not; if datum is not easily available, change wording to near-original 'faces with 0--2 wings.' Flowering summer (Jul--Sep). Slightly basic to slightly acidic to alkaline waters of ponds, lakes, and swamps; 100---1800 m; Sask.; Ala., Ark., Calif., Colo., Ill., Ind., Iowa, Kans., Ky., La., Mich., Minn., Miss., Mo., Nebr., Ohio, Okla., S.Dak., Tex., Va., Wis.
Perennial aquatic herb to 0.7 m tall Leaves: emersed, with a ridged stalk to 39 cm long and an arrowhead-shaped blade 5 - 20 cm long and 2 - 8 cm wide. The leaf lobes are shorter than or equal in length to the rest of the blade. Inflorescence: emersed, singly stalked along a central axis (raceme), or with loosely branched stalks (panicle), 25 - 30 cm long, 6 - 15 cm wide, arising from a main stalk to 0.8 m long, with five to twelve whorls of flowers. Flowers: either male or female, found on the same plant (monoecious), to 3.5 cm wide, with three recurved to spreading sepals and three white petals 8 - 12 mm long. Male flowers are borne on the upper part of the inflorescence, having smooth cylindric filaments longer than the anthers. Female flowers borne on lower part of the inflorescence. Fruit: a spherical cluster of achenes 1.5 - 2.5 cm wide. Each achene is 2 - 3 mm long, 1.4 - 2.2 mm wide, wedge-shaped to inversely egg-shaped, minutely shallow-toothed, with an ascending beak 0.4 - 1.7 mm long. Bracts: straw-colored, 1 - 4 cm long, linear to lance-shaped, pointed, firm, papery, separate or fused to one-quarter the length.
Similar species: Sagittaria latifolia, Sagittaria brevirostra, Sagittaria cuneata, and Sagittaria montevidensis ssp. calycina all have emersed arrowhead-shaped leaves and filaments lacking scales. Sagittaria latifolia has nearly all emersed leaves that are highly variable (linear to arrowhead-shaped) and achenes with beaks spreading horizontally. Sagittaria cuneata has submersed ribbon-like leaves, floating lance- to arrowhead-shaped leaves, emersed arrowhead-shaped leaves, and achenes with short, erect beaks. Sagittaria montevidensis ssp. calycina has linear submersed leaves, arrowhead-shaped emersed leaves, and achenes with horizontal beaks. While the other species above have separate male and female flowers and reflexed or spreading sepals, S. montevidensis ssp. calycina has male and bisexual flowers. The bisexual flowers are distinguished by a ring of sterile stamens and appressed sepals.
Flowering: early July to late August
Habitat and ecology: Occasional in shallow water, marshes, sloughs, and muddy marshy edges of ponds and streams.
Occurence in the Chicago region: native
Etymology: Sagittaria comes from the Latin word sagitta, meaning arrow, referring to the leaf shape. Brevirostra means short-beaked.
Author: The Morton Arboretum
Main portion of the lf-blade mostly lanceolate, sometimes broader, mostly 5-20 נ2-8 cm, 0.8-8 times as long as wide; petiole corrugated in x-section, at least in the lower half; infl often branched at the lowest whorl; fls in 5-12 whorls; bracts 1.5-4 cm, equaling or longer than the 1.3-2.5 cm pedicels, stramineous, papery-firm, broad-based and long-attenuate; fruiting head usually oblate-spheroidal; receptacle not markedly echinate; achenes 2-3 mm, lacking resin-ducts, with a short wing (not extending into the beak) on one or both faces, or the faces wingless; dorsal wing entire to crenulate, usually truncate at the top and not confluent with the beak; ventral wing narrower and confluent with the beak; beak 0.4-1.7 mm; 2n=22; otherwise much like no. 4 [Sagittaria engelmanniana J. G. Sm]. In circumneutral to somewhat alkaline shallow water of ponds and swamps; O. and Mich. to Minn. and S.D., s. to Tenn., Ala., and Tex. July-Oct. (S. engelmanniana var. b.)
Gleason, Henry A. & Cronquist, Arthur J. 1991. Manual of vascular plants of northeastern United States and adjacent Canada. lxxv + 910 pp.
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